View source for Social:Archaeogenetics of the Near East

The '''archaeogenetics of the Near East''' is the study of the genetics of past human populations (archaeogenetics) in the Ancient Near East using [[Biology:DNA|DNA]] from ancient remains. Researchers use Y-DNA, mtDNA, and other autosomal DNAs to identify haplogroups and haplotypes in ancient populations of Egypt, Persia, Mesopotamia, Anatolia, Arabia, the Levant, and other areas.{{fact|date=June 2019}}

==History==
Developments in DNA sequencing in the 1970s and 1980s provided researchers with the tools needed to study human genetic variation and the genetics of human populations to discover founder populations of modern people groups and human migrations.{{fact|date=June 2019}}

In 2005, National Geographic launched The Genographic Project, led by 12 prominent scientists and researchers, to study and map historical human migration patterns by collecting and analyzing [[Biology:DNA|DNA]] samples from hundreds of thousands of people from around the world.{{fact|date=June 2019}}

==Egypt==
{{main|DNA history of Egypt}}
Contamination from handling and intrusion from microbes create obstacles to the recovery of [[Biology:Ancient DNA|Ancient DNA]].<ref name="books.google.com">{{cite book|author1=Kathryn A. Bard|author2=Steven Blake Shubert|title=Encyclopedia of the Archaeology of Ancient Egypt|url=https://books.google.com/books?id=XNdgScxtirYC|year=1999|publisher=Taylor & Francis|isbn=978-0-203-98283-9|pages=278–279}}</ref> Consequently, most DNA studies have been carried out on modern Egyptian populations with the intent of learning about the influences of historical migrations on the population of Egypt.<ref name="S.O.Y. Keita & A. J. Boyce">{{cite journal |doi=10.1353/hia.2005.0013 |jstor=20065742 |title=Genetics, Egypt, and History: Interpreting Geographical Patterns of Y Chromosome Variation |journal=History in Africa |volume=32 |pages=221–46 |date=2005 |last1=Keita |first1=S. O. Y. |last2=Boyce |first2=A. J.}}</ref><ref name="Shomarka Keita (2005)">{{cite journal |doi=10.1007/s10437-005-4189-4 |jstor=25130819 |title=Explanation of the Pattern of P49a,f ''Taq''I RFLP Y-Chromosome Variation in Egypt |journal=African Archaeological Review |volume=22 |issue=2 |pages=61–75 |date=2005 |last1=Keita |first1=S. O. Y.}}</ref><ref name="Keita">{{cite journal |author=Keita SO |title=History in the interpretation of the pattern of p49a,f TaqI RFLP Y-chromosome variation in Egypt: a consideration of multiple lines of evidence |pages=559–67  |journal=American Journal of Human Biology |volume=17 |issue=5 |date=2005 |pmid=16136533 |doi=10.1002/ajhb.20428}}</ref><ref>{{cite web|url=http://ngm.nationalgeographic.com/video/player?titleID=1414281487#/?titleID=1414242821&catID=1 |title=Shomarka Keita: What genetics can tell us |publisher=Ngm.nationalgeographic.com |accessdate=2014-06-30}}</ref>

In general, various DNA studies have found that the gene frequencies of North African populations are intermediate between those of the Near East, the Horn of Africa, southern Europe and Sub Saharan Africa,<ref>[https://books.google.com/books?id=FrwNcwKaUKoC&printsec=frontcover#PPA136,M1 Cavalli-Sforza, History and Geography of Human Genes], The intermediacy of North Africa and to a lesser extent Europe is apparent</ref> though Egypt's NRY frequency distributions appear to be much more similar to those of the Middle East than to any sub-Saharan African population, suggesting a much larger Eurasian genetic component.<ref name="luis">{{cite journal |last1=Luis |first1=J |last2=Rowold |first2=D |last3=Regueiro |first3=M |last4=Caeiro |first4=B |last5=Cinnioglu |first5=C |last6=Roseman |first6=C |last7=Underhill |first7=P |last8=Cavallisforza |first8=L |last9=Herrera |first9=R  |date=2004 |title=The Levant versus the Horn of Africa: Evidence for Bidirectional Corridors of Human Migrations |journal=The American Journal of Human Genetics |volume=74 |issue=3 |pages=532–44 |pmid=14973781 |pmc=1182266 |doi=10.1086/382286}}</ref><ref name="Cavalli-Sforza">Cavalli-Sforza, L.L., P. Menozzi, and A. Piazza. 1994. The History and Geography of Human Genes. Princeton: Princeton University Press.</ref><ref name="Cavalli-Sforza" /><ref name="Bosch1997">{{cite journal |author=Arredi B |title=A predominantly neolithic origin for Y-chromosomal DNA variation in North Africa |journal=American Journal of Human Genetics |volume=75 |issue=2 |pages=338–45 |date=August 2004  |pmid=15202071 |pmc=1216069 |doi=10.1086/423147|name-list-format=vanc |author2= Poloni ES |author3= Paracchini S  |displayauthors=etal  }}</ref><ref>{{cite journal |author=Manni F |title=Y-chromosome analysis in Egypt suggests a genetic regional continuity in Northeastern Africa |journal=Human Biology |volume=74 |issue=5 |pages=645–58 |date=October 2002  |pmid=12495079 |doi=10.1353/hub.2002.0054 |url=http://digitalcommons.wayne.edu/humbiol/vol74/iss5/2/|name-list-format=vanc |author2= Leonardi P |author3= Barakat A  |displayauthors=etal  }}</ref><ref>{{cite book|title= The History and Geography of Human Genes|last=Cavalli-Sforza|chapter=Synthetic maps of Africa|chapterurl=https://books.google.com/books?id=FrwNcwKaUKoC&printsec=frontcover#PPA189,M1|isbn= 978-0691087504|year=1994}}The present population of the Sahara is Sudan in the extreme north, with an increase of Negroid component as one goes south</ref>

Blood typing and DNA sampling on ancient Egyptian mummies is scant; however, a 1982 study of blood typing of dynastic mummies found ABO frequencies to be most similar to modern Egyptians<ref>{{cite journal |id={{INIST|12409492}} |last1=Borgognini Tarli |first1=S.M. |last2=Paoli |first2=G. |date=1982 |title=Survey on paleoserological studies |journal=Homo Gottingen |volume=33 |issue=2–3 |pages=69–89}}</ref> and some also to Northern Haratin populations. ABO blood group distribution shows that the Egyptians form a sister group to North African populations, including Berbers, Nubians and Canary Islanders.<ref>{{cite book |last1=Cavalli-Sforza |first1=L.L. |first2=P. |last2=Menozzi |first3=A. |last3=Piazza |date=1994 |title=The History and Geography of Human Genes |location=Princeton |publisher=Princeton University Press |pages=169–74}}</ref>

In 2013, ''Nature'' announced the publication of the first genetic study utilizing [[Biology:Whole genome sequencing|next-generation sequencing]] to ascertain the ancestral lineage of an Ancient Egyptian individual. The research was led by Carsten Pusch of the University of Tübingen in Germany and Rabab Khairat, who released their findings in the ''Journal of Applied Genetics''. DNA was extracted from the heads of five Egyptian mummies that were housed at the institution. All the specimens were dated between 806 BC and 124 AD, a timeframe corresponding with the late Dynastic period. The researchers observed that one of the mummified individuals likely belonged to the mtDNA haplogroup I2, a maternal clade that is believed to have originated in Western Asia.<ref name=Marchant>{{cite news|last=Marchant|first=Jo|title=Egyptian mummies yield genetic secrets|url=http://www.nature.com/news/egyptian-mummies-yield-genetic-secrets-1.12793#/b1|accessdate=13 January 2014|newspaper=Nature|date=12 April 2013}}</ref>

== Iran (Persia) ==

A recent study analyzed the autosomal DNA and genome of an Iron Age Iranian sample taken from Teppe Hasanlu (F38_Hasanlu, '''dated to 971-832 BCE''') and revealed it has close affinities to modern Iranians.<ref name="ReferenceA">{{cite journal | last1 = Broushaki | first1 = F. | last2 = Thomas | first2 = M. | last3 = Link | first3 = V. |display-authors=et al | year = 2016 | title = Early Neolithic genomes from the eastern Fertile Crescent | journal = Science | volume = 353 | issue = 6298 |pages = 499–503 | doi=10.1126/science.aaf7943| pmid = 27417496 | pmc = 5113750 | bibcode = 2016Sci...353..499B }}</ref> The sample's data exists on GEDMatch under the kit number M381564. This indicates strong genetic continuity between modern Iranians and their Iron Age ancestors, discrediting Nordicist claims, such as those made by Arthur de Gobineau, that the ancient Achaemenids were genetically Western European.

The latest comparative study (2013) on the complete mitochondrial DNA diversity in Iranians has indicated that Iranian Azeris are more related to the people of Georgia, than they are to other Iranians (Persians, Armenians, etc.), while the Persians, Armenians and Qashqai on the other hand were more related to each other.<ref name="ReferenceB">{{cite journal | last1 = Derenko | first1 = M. | last2 = Malyarchuk | first2 = B. | last3 = Bahmanimehr | first3 = A. | last4 = Denisova | first4 = G. | last5 = Perkova | first5 = M. | last6 = Farjadian | first6 = S. | last7 = Yepiskoposyan | first7 = L. | year = 2013 | title = Complete Mitochondrial DNA Diversity in Iranians | journal = PLoS ONE | volume = 8 | issue = 11| page = e80673 | doi=10.1371/journal.pone.0080673 | pmid=24244704 | pmc=3828245| bibcode = 2013PLoSO...880673D }}</ref>

It furthermore showed that overall, the complete mtDNA sequence analysis revealed an extremely high level of genetic diversity in the Iranian populations studied which is comparable to the other groups from the South Caucasus, Anatolia and Europe.<ref name="ReferenceB"/> The same 2013 research further noted that "the results of AMOVA and MDS analyses did not associate any regional and/or linguistic group of populations in the Anatolia, Caucasus and Iran region pointing to strong genetic affinity of Indo-European speaking Persians and Turkic-speaking Qashqais, thus suggesting their origin from a common maternal ancestral gene pool.<ref name="ReferenceB"/> The pronounced influence of the South Caucasus populations on the maternal diversity of Iranian Azeris is also evident from the MDS analysis results."<ref name="ReferenceB"/>

The study also notes that " It is worth pointing out the position of Azeris from the Caucasus region, who despite their supposed common origin with Iranian Azeris, cluster quite separately and occupy an intermediate position between the Azeris/Georgians and Turks/Iranians grouping".<ref name="ReferenceB"/>

The MtDNA results from the samples overall on average closely resemble those found in the neighbouring regions of the Caucasus, Anatolia, and to a lesser extent (Northern) Mesopotamia.<ref name="ReferenceB"/>

Among the most common MtDNA lineages in the nation, namely U3b3, appears to be restricted to populations of Iran and the Caucasus, while the sub-cluster U3b1a is common in the whole Near East region.<ref name="plosone.org">{{cite journal|title=Complete Mitochondrial DNA Diversity in Iranians|journal=PLOS ONE|volume=8|issue=11|pages=e80673|doi=10.1371/journal.pone.0080673|pmid=24244704|pmc=3828245|year = 2013|last1 = Derenko|first1 = Miroslava|last2=Malyarchuk|first2=Boris|last3=Bahmanimehr|first3=Ardeshir|last4=Denisova|first4=Galina|last5=Perkova|first5=Maria|last6=Farjadian|first6=Shirin|last7=Yepiskoposyan|first7=Levon}}</ref>

An earlier genetic research was made by ''Nasidze et al.'' (2006) on the North Iranian populations on the Gilaks and Mazandaranis, spanning the southwestern coast of the Caspian Sea, up to the border with neighbouring Azerbaijan. The Gilaks and Mazandaranis comprise 7% of the Iranian population. It has been suggested that their ancestors came from the Caucasus region, perhaps displacing an earlier group in the South Caspian.<ref name="sciencedirect.com">{{cite journal|title=''Concomitant Replacement of Language and mtDNA in South Caspian Populations of Iran''| doi=10.1016/j.cub.2006.02.021|pmid=16581511|volume=16|issue=7|date=April 2006|journal=Curr. Biol.|pages=668–73 | last1 = Nasidze | first1 = I | last2 = Quinque | first2 = D | last3 = Rahmani | first3 = M | last4 = Alemohamad | first4 = SA | last5 = Stoneking | first5 = M}}</ref> Linguistic evidence supports this scenario, in that the Gilaki and Mazandarani languages (but not other Iranian languages) share certain typological features with Caucasian languages, and specifically South Caucasian languages.<ref name="sciencedirect.com"/> There have been patterns analyzed of mtDNA and Y chromosome variation in the Gilaki and Mazandarani.

Based on mtDNA HV1 sequences tested by ''Nasidze et al''., the Gilaks and Mazandarani most closely resemble their geographic and linguistic neighbors, namely other Iranian groups. However, their Y chromosome types most closely resemble those found in groups from the South Caucasus.<ref name="sciencedirect.com"/>
A scenario that explains these differences is a south Caucasian origin for the ancestors of the Gilani and Mazandarani, followed by introgression of women (but not men) from local Iranian groups, possibly because of patrilocality.<ref name="sciencedirect.com"/>
Given that both mtDNA and language are maternally transmitted, the incorporation of local Iranian women would have resulted in the concomitant replacement of the ancestral Caucasian language and mtDNA types of the Gilani and Mazandarani with their current Iranian language and mtDNA types. Concomitant replacement of language and mtDNA may be a more general phenomenon than previously recognized.

The Mazandarani and Gilani groups fall inside a major cluster consisting of populations from the Caucasus and West Asia and are particularly close to the South Caucasus groups—Georgians, Armenians, and Azerbaijanis. Iranians from Tehran and Isfahan are situated more distantly from these groups.<ref name="sciencedirect.com"/>

==Iraq (Mesopotamia)==

In the 1995 book ''The History and Geography of Human Genes'' the authors wrote that: "The Assyrians are a fairly homogeneous group of people, believed to originate from the land of old Assyria in northern Iraq [..] they are Christians and are bona fide descendants of their ancient namesakes."<ref>Luigi Luca Cavalli-Sforza, Paolo Menozzi, Alberto Piazza, The History and Geography of Human Genes, p. 243</ref>

In a 2006 study of the Y chromosome DNA of six regional populations, including, for comparison, Assyrians and Syrians, researchers found that, "the two Semitic populations (Assyrians and Syrians) are very distinct from each other according to both [comparative] axes. This difference supported also by other methods of comparison points out the weak genetic affinity between the two populations with different historical destinies." <ref name="Iran and the Caucasus">[http://www.rau.am/downloads/publ.kafedr/episkoposyan_medbiolog/Yepiskoposian_I&C_06.pdf Yepiskoposian et al., Iran and the Caucasus, Volume 10, Number 2, 2006, pp. 191–208(18), "Genetic Testing of Language Replacement Hypothesis in Southwest Asia"]</ref>

A 2008 study on the genetics of "old ethnic groups in Mesopotamia," including 340 subjects from seven ethnic communities ("These populations included Assyrians, Jews, Zoroastrians, Armenians, and Turkmen (representing ethnic groups from Iran, restricted by rules of their religion), and the Iraqi and Kuwaiti populations from Iraq and Kuwait.") found that Assyrians were homogeneous with respect to all other ethnic groups sampled in the study, regardless of religious affiliation.<ref name="pubmed.gov">{{cite journal | pmid = 18505046 | doi=10.3378/1534-6617(2008)80[73:VODVAA]2.0.CO;2 | volume=80 | title=Variation of DAT1 VNTR alleles and genotypes among old ethnic groups in Mesopotamia to the Oxus region | year=2008 | journal=Hum Biol | pages=73–81 | last1 = Banoei | first1 = MM | last2 = Chaleshtori | first2 = MH | last3 = Sanati | first3 = MH | last4 = Shariati | first4 = P | last5 = Houshmand | first5 = M | last6 = Majidizadeh | first6 = T | last7 = Soltani | first7 = NJ | last8 = Golalipour | first8 = M}} "The relationship probability was lowest between Assyrians and other communities. Endogamy was found to be high for this population through determination of the heterogeneity coefficient (+0,6867), Our study supports earlier findings indicating the relatively closed nature of the Assyrian community as a whole, which as a result of their religious and cultural traditions, have had little intermixture with other populations."</ref>

A study published in 2011 looking at the relationship between Iraq's Marsh Arabs and ancient [[Social:Sumer|Sumer]]ians concluded "the modern Marsh Arabs of Iraq harbour mtDNAs and Y chromosomes that are predominantly of Middle Eastern origin. Therefore, certain cultural features of the area such as water buffalo breeding and rice farming, which were most likely introduced from the Indian sub-continent, only marginally affected the gene pool of the autochthonous people of the region. Moreover, a Middle Eastern ancestral origin of the modern population of the marshes of southern Iraq implies that, ''if'' the Marsh Arabs are descendants of the ancient Sumerians, also Sumerians were not of Indian or Southern Asian ancestry."<ref name=BMC>{{cite journal |author=Al-Zahery N |title=In search of the genetic footprints of Sumerians: a survey of Y-chromosome and mtDNA variation in the Marsh Arabs of Iraq |journal=BMC Evolutionary Biology |volume=11 |page=288 |date=2011 |pmid=21970613 |pmc=3215667 |doi=10.1186/1471-2148-11-288|name-list-format=vanc |author2= Pala M |author3= Battaglia V |displayauthors=3 |last4=Grugni  | first4=V. |last5=Hamod |first5=Mohammed A |last6=Kashani |first6=Baharak |last7=Olivieri |first7=Anna |last8=Torroni |first8=Antonio |last9=Santachiara-Benerecetti |first9=Augusta S |last10=Semino |first10=Ornella }}</ref>

In a 2011 study focusing on the genetics of the Maʻdān people of Iraq, researchers identified Y chromosome haplotypes shared by Marsh Arabs, Arabic speaking Iraqis, Assyrians and Mandeans  "supporting a common local background."<ref name=BMC/>

A 2013 study based on DNA extracted from the dental remains of four individuals, unearthed at Tell Ashara (ancient Terqa) and Tell Masaikh (ancient Kar-Assurnasirpal) suggest a genetic link between the people of Bronze Age Mesopotamia and South Asia. According to the study, "We anticipate that the analysed remains from Mesopotamia belonged to people with genetic affinity to the Indian subcontinent since the distribution of identified ancient haplotypes indicates solid link with populations from the region of South Asia-Tibet (Trans-Himalaya). They may have been descendants of migrants from much earlier times, spreading the clades of the macrohaplogroup M throughout Eurasia and founding regional Mesopotamian groups like that of Terqa or just merchants moving along trade routes passing near or through the region."<ref>{{cite journal | last1 = Witas | first1 = HW | last2 = Tomczyk | first2 = J | last3 = Jędrychowska-Dańska | first3 = K | last4 = Chaubey | first4 = G | last5 = Płoszaj | first5 = T | year = 2013 | title = mtDNA from the Early Bronze Age to the Roman Period Suggests a Genetic Link between the Indian Subcontinent and Mesopotamian Cradle of Civilization | journal = PLoS ONE | volume = 8 | issue = 9| page = e73682 | doi = 10.1371/journal.pone.0073682 | pmid=24040024 | pmc=3770703| bibcode = 2013PLoSO...873682W }}</ref> A 2014 study expanding on the 2013 study and based on analysis of 15751 DNA samples arrives at the conclusion that the sources for South Asian DNA in Mesopotamian populations are Tamil merchants involved in trade with Rome, stating "M65a, M49 and/or M61 haplogroups carrying ancient Mesopotamians might have been the merchants from India".<ref>{{cite journal | last1 = Palanichamy Mg | first1 = Mitra B | last2 = Debnath | first2 = M | last3 = Agrawal | first3 = S | last4 = Chaudhuri | first4 = TK | last5 = Zhang | first5 = Y-P | year = 2014 | title = Tamil Merchant in Ancient Mesopotamia | url = | journal = PLoS ONE | volume = 9 | issue = 10| page = e109331 | doi = 10.1371/journal.pone.0109331 | pmid = 25299580 | pmc = 4192148 | bibcode = 2014PLoSO...9j9331P }}</ref>

=={{anchor|Levant}}Levant (Israel, Syria, Palestine, Lebanon, Jordan)==

Zalloua and Wells (2004), under the auspices of a  grant from National Geographic Magazine, examined the origins of the Canaanite Phoenicians.
The debate between Wells and Zalloua was whether haplogroup J2 (M172) should be identified as that of the Phoenicians or that of its "parent" haplogroup M89 on the YDNA phylogenetic tree.<ref>{{cite web|url=http://ycc.biosci.arizona.edu/nomenclature_system/fig1.html |archive-url=https://archive.is/20120805220306/http://ycc.biosci.arizona.edu/nomenclature_system/fig1.html |dead-url=yes |archive-date=August 5, 2012 |accessdate=September 16, 2007 }}</ref> 
Initial consensus suggested that J2 be identified with the Canaanite-Phoenician (Northwest Semitic) population, with avenues open for future research.<ref>National Geographic Magazine, October 2004. Available online: http://ngm.nationalgeographic.com/features/world/asia/lebanon/phoenicians-text/1; and http://www.independent.com.mt/news.asp?newsitemid=57215 [accessed: March 10, 2008]</ref> As Wells commented, "The Phoenicians were the Canaanites"<ref>{{cite web|url=http://ngm.nationalgeographic.com/features/world/asia/lebanon/phoenicians-text/5 |title=Who Were the Phoenicians? - National Geographic Magazine |publisher=Ngm.nationalgeographic.com |accessdate=2014-06-30}}</ref> 
It was reported in the PBS description of the National Geographic TV Special on this study entitled "Quest for the Phoenicians" that ancient DNA was included in this study as extracted from the tooth of a 2500-year-old Phoenician mummy.<ref>{{cite web|url=https://www.pbs.org/previews/phoenicians/|year=2004 |title=National Geographic Special 'Quest for the Phoenicians' |publisher=PBS |archiveurl=https://web.archive.org/web/20040923131922/http://www.pbs.org/previews/phoenicians/|archivedate=2004-09-23}}</ref>

Wells identified the haplogroup of the Canaanites as haplogroup J2 which originated in northern Mesopotamia.<ref>[http://ngm.nationalgeographic.com/features/world/asia/lebanon/phoenicians-text/5; and http://www.independent.com.mt/news.asp?newsitemid=57215 [Accessed April 11, 2008] </ref> The National Geographic Genographic Project linked haplogroup J2 to the site of Jericho, Tel el-Sultan, ca. 8500 BCE and indicated that in modern populations, haplogroup J2 is found primarily in the Middle East, but also along the coasts of North Africa and Southern Europe, with especially high distribution among present-day Jewish populations (30%), Southern Italians (20%), and lower frequencies in Southern Spain (10%).<ref>The Atlas of the Human Journey-Genetic Markers-Haplogroup J2 (M172): {{cite web|url=https://www3.nationalgeographic.com/genographic/atlas.html |title=Archived copy |accessdate=2013-03-26 |deadurl=yes |archiveurl=https://web.archive.org/web/20080405232839/https://www3.nationalgeographic.com/genographic/atlas.html |archivedate=2008-04-05 |df= }} [Accessed April 11, 2008]</ref>

In a 2005 study of ASPM gene variants, Mekel-Bobrov et al. found that the Israeli Druze people of the Carmel region have among the highest rate of the newly evolved ASPM haplogroup D, at 52.2% occurrence of the approximately 6,000-year-old allele.<ref>{{cite journal |author=Mekel-Bobrov N |title=Ongoing adaptive evolution of ASPM, a brain size determinant in Homo sapiens |journal=Science |volume=309 |issue=5741 |pages=1720–2 |date=September 2005  |pmid=16151010 |doi=10.1126/science.1116815 |bibcode=2005Sci...309.1720M|name-list-format=vanc |author2= Gilbert SL |author3= Evans PD |displayauthors=3 |last4=Vallender|first4=Eric J. |last5=Anderson |first5=Jeffrey R. |last6=Hudson |first6=Richard R. |last7=Tishkoff |first7=Sarah A. |last8=Lahn |first8=Bruce T. }}</ref> While it is not yet known exactly what selective advantage is provided by this gene variant, the haplogroup D allele is thought to be positively selected in populations and to confer some substantial advantage that has caused its frequency to rapidly increase.
According to [[Biology:DNA|DNA]] testing, Druze are remarkable for the high frequency (35%) of males who carry the Y-chromosomal haplogroup L, which is otherwise uncommon in the Mideast (Shen et al. 2004).<ref name="Shen P 2004 248–60">{{cite journal |author=Shen P |title=Reconstruction of patrilineages and matrilineages of Samaritans and other Israeli populations from Y-chromosome and mitochondrial DNA sequence variation |journal=Human Mutation |volume=24 |issue=3 |pages=248–60 |date=September 2004  |pmid=15300852 |doi=10.1002/humu.20077|name-list-format=vanc |author2= Lavi T |author3= Kivisild T  |displayauthors=etal  }}</ref> This haplogroup originates from prehistoric South Asia and has spread from Pakistan into southern Iran.

Cruciani in 2007 found E1b1b1a2 (E-V13) [one from Sub Clades of E1b1b1a1 (E-V12)] in high levels (>10% of the male population) in Turkish Cypriot and Druze Arab lineages. Recent genetic clustering analyses of ethnic groups are consistent with the close ancestral relationship between the Druze and Cypriots, and also identified similarity to the general Syrian and Lebanese populations, as well as a variety of Jewish lineages (Ashkenazi, Sephardi, Iraqi Jewish, and Moroccan Jews) (Behar et al. 2010).<ref name="Behar2010">{{cite journal |author=Behar DM |title=The genome-wide structure of the Jewish people |journal=Nature |volume=466 |issue=7303 |pages=238–42 |date=July 2010  |pmid=20531471 |doi=10.1038/nature09103|name-list-format=vanc |author2= Yunusbayev B |author3= Metspalu M |author4=Metspalu, Ene |displayauthors=3 |bibcode=2010Natur.466..238B |last5=Rosset |first5=Saharon |last6=Parik |first6=Jüri |last7=Rootsi |first7=Siiri |last8=Chaubey |first8=Gyaneshwer |last9=Kutuev |first9=Ildus |last10=Yudkovsky |first10=Guennady |last11=Khusnutdinova |first11=Elza K. |last12=Balanovsky |first12=Oleg |last13=Semino |first13=Ornella |last14=Pereira |first14=Luisa |last15=Comas |first15=David |last16=Gurwitz |first16=David |last17=Bonne-Tamir |first17=Batsheva |last18=Parfitt |first18=Tudor |last19=Hammer |first19=Michael F. |last20=Skorecki |first20=Karl |last21=Villems |first21=Richard }}</ref>

A study published by the National Academy of Sciences found that "the [[Biology:Y chromosome|paternal gene]] pools of Jewish communities from Europe, North Africa, and the Middle East descended from a common Middle Eastern ancestral population", and suggested that "most Jewish communities have remained relatively isolated from neighbouring non-Jewish communities during and after the Diaspora".<ref name="pmid10801975">{{cite journal |author=Hammer MF |title=Jewish and Middle Eastern non-Jewish populations share a common pool of Y-chromosome biallelic haplotypes |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=97 |issue=12 |pages=6769–74 |date=June 2000  |pmid=10801975 |pmc=18733 |doi=10.1073/pnas.100115997 |bibcode=2000PNAS...97.6769H|name-list-format=vanc |author2= Redd AJ |author3= Wood ET |displayauthors=3 |author4=Bonner, M. R. |last5=Jarjanazi |first5=H. |last6=Karafet |first6=T. |last7=Santachiara-Benerecetti |first7=S. |last8=Oppenheim |first8=A. |last9=Jobling |first9=M. A. |last10=Jenkins |first10=T. |last11=Ostrer |first11=H. |last12=Bonné-Tamir |first12=B. }}</ref> Researchers expressed surprise at the remarkable genetic uniformity they found among modern Jews, no matter where the diaspora has become dispersed around the world.<ref name="pmid10801975"/>
Skorecki and colleague wrote that "the extremely close affinity of Jewish and non-Jewish Middle Eastern populations observed ... supports the hypothesis of a common Middle Eastern origin".<ref name="hammer2">{{cite journal|author=Skorecki K |title=Y chromosomes of Jewish priests |journal=Nature |volume=385 |issue=6611 |page=32 |date=January 1997 |pmid=8985243 |doi=10.1038/385032a0 |url=http://www.familytreedna.com/nature97385.html |bibcode=1997Natur.385...32S |author2=Selig S |author3=Blazer S |displayauthors=etal |deadurl=yes |archiveurl=https://web.archive.org/web/20070209230313/http://www.familytreedna.com/nature97385.html |archivedate=February 9, 2007 }}</ref>

This research has suggested that, in addition to Israelite male, significant female founder ancestry might also derive from the Middle East-with 40% of Ashkenazim descended from four women who lived about 2000–3000 years ago in the Middle East.<ref name="wade">{{Cite news|url=https://www.nytimes.com/2006/01/14/science/14gene.html?_r=1&oref=slogin|  title=New Light on Origins of Ashkenazi in Europe|first=Nicholas|last=Wade|journal=The New York Times|date=January 14, 2006|accessdate=24 May 2006}}</ref> In addition, Behar (2006) suggested that the rest of Ashkenazi mtDNA is originated from about 150 women; most of those were probably of Middle Eastern origin.<ref name="behar2006">{{cite journal |author=Behar DM |title=The matrilineal ancestry of Ashkenazi Jewry: portrait of a recent founder event |journal=American Journal of Human Genetics |volume=78 |issue=3 |pages=487–97 |date=March 2006  |pmid=16404693 |pmc=1380291 |doi=10.1086/500307|name-list-format=vanc |author2= Metspalu E |author3= Kivisild T  |displayauthors=etal  }}</ref>  A 2013 genetic study suggested that the four founding maternal lineages of Ashkenazi Jews originate in Europe and that only ~8% of Ashkenazi mtDNA can confidently be assigned a Near Eastern origin, while >80% of Ashkenazi maternal lineages have a likely European origin <ref name="costa2013">{{cite journal |author=Costa MD |title=A substantial prehistoric European ancestry amongst Ashkenazi maternal lineages |journal=Nature Communications |volume=4 |page=2543 |date=2013 |pmid=24104924 |pmc=3806353 |doi=10.1038/ncomms3543|name-list-format=vanc |author2= Pereira JB |author3= Pala M |displayauthors=3 |first4=Verónica |last4=Fernandes |last5=Olivieri |first5=Anna |last6=Achilli |first6=Alessandro |last7=Perego |first7=Ugo A. |last8=Rychkov |first8=Sergei |last9=Naumova |first9=Oksana |last10=Hatina |first10=Jiři |last11=Woodward |first11=Scott R. |last12=Eng |first12=Ken Khong |last13=MacAulay |first13=Vincent |last14=Carr |first14=Martin |last15=Soares |first15=Pedro |last16=Pereira |first16=Luísa |last17=Richards |first17=Martin B. |bibcode = 2013NatCo...4E2543C }}</ref> while the 2014 study carried out by Spanish geneticists suggested ancient Near Eastern origin of four founding maternal lineages of Ashkenazi Jews.<ref>{{Cite journal | doi=10.1371/journal.pgen.1004401| pmid=24901650| pmc=4046922|title = Ancient DNA Analysis of 8000 B.C. Near Eastern Farmers Supports an Early Neolithic Pioneer Maritime Colonization of Mainland Europe through Cyprus and the Aegean Islands| journal=PLOS Genetics| volume=10| issue=6| pages=e1004401|year = 2014|last1 = Fernández|first1 = Eva| last2=Pérez-Pérez| first2=Alejandro| last3=Gamba| first3=Cristina| last4=Prats| first4=Eva| last5=Cuesta| first5=Pedro| last6=Anfruns| first6=Josep| last7=Molist| first7=Miquel| last8=Arroyo-Pardo| first8=Eduardo| last9=Turbón| first9=Daniel}}</ref>

In 2004, a team of geneticists from Stanford University, the Hebrew University of Jerusalem, Tartu University (Estonia), Barzilai Medical Center (Ashkelon, Israel), and the Assaf Harofeh Medical Center (Zerifin, Israel), studied the modern Samaritan ethnic community living in Israel in comparison with modern Israeli populations to explore the ancient genetic history of these people groups. The Samaritans or ''Shomronim'' (singular: ''Shomroni''; Hebrew: שומרוני) trace their origins to the Assyrian province of Shomron (Samaria) in ancient Israel in the period after the Assyrian conquest circa 722 BCE. Shomron was the capital of the Northern Kingdom of Israel when it was conquered by the Assyrians and gave the name to the ancient province of Samaria and the Samaritan people group. Tradition holds that the Samaritans were a mixed group of Israelites who were not exiled or were sent back or returned from exile and non-Israelites relocated to the region by the Assyrians. The modern-day Samaritans are believed to be the direct descendants of the ancient Samaritans.

Their findings reported on four family lineages among the Samaritans: the Tsdaka family  (tradition: tribe of Menasseh), the Joshua-Marhiv and Danfi families (tradition: tribe of Ephraim), and the Cohen family (tradition: tribe of Levi). All Samaritan families were found in haplogroups J1 and J2, except the Cohen family which was found in haplogroup E3b1a-M78.<ref name="Shen P 2004 248–60"/> This article predated the E3b1a subclades based on the research of Cruciani, et al.<ref>{{cite journal |author=Cruciani F |author2=La Fratta R |author3=Torroni A |author4=Underhill PA |author5=Scozzari R |title=Molecular dissection of the Y chromosome haplogroup E-M78 (E3b1a): a posteriori evaluation of a microsatellite-network-based approach through six new biallelic markers |journal=Human Mutation |volume=27 |issue=8 |pages=831–2 |date=August 2006  |pmid=16835895 |doi=10.1002/humu.9445}}</ref>

A 2018 study conducted by scholars from Tel-Aviv University, the [[Social:Israel Antiquities Authority|Israel Antiquities Authority]] and Harvard University had discovered that 22 out of the 600 people who were buried in Peki'in cave from the [[Chemistry:Chalcolithic|Chalcolithic]] Period were of both local Levantine and Persian and Zagros<ref>[https://www.jpost.com/Israel-News/DNA-study-in-Israeli-cave-sheds-light-on-origins-of-Chalcolithic-culture-565293 DNA STUDY IN ISRAELI CAVE SHEDS LIGHT ON ORIGINS OF CHALCOLITHIC CULTURE]</ref> area ancestries, or as phrased in the paper itself: "Ancient DNA from Chalcolithic Israel reveals the role of population mixture in cultural transformation,” the scientists concluded that the homogeneous community found in the cave could source ~57% of its ancestry from groups related to those of the local Levant Neolithic, ~26% from groups related to those of the Anatolian Neolithic, and ~17% from groups related to those of the Iran Chalcolithic.".<ref name="timesofisrael.com">[https://www.timesofisrael.com/anomalous-blue-eyed-people-came-to-israel-6500-years-ago-from-iran-dna-shows/ Anomalous blue-eyed people came to Israel 6,500 years ago from Iran, DNA shows]</ref> The scholars noted that the Zagros genetic material held “Certain characteristics, such as genetic mutations contributing to blue eye color, were not seen in the DNA test results of earlier Levantine human remains...The blue-eyed, fair-skinned community didn’t continue, but at least now researchers have an idea why. “These findings suggest that the rise and fall of the Chalcolithic culture are probably due to demographic changes in the region”.<ref name="timesofisrael.com"/><ref>{{Cite journal | doi=10.1038/s41467-018-05649-9| pmid=30127404| pmc=6102297| title=Ancient DNA from Chalcolithic Israel reveals the role of population mixture in cultural transformation| journal=Nature Communications| volume=9| issue=1| pages=3336| year=2018| last1=Harney| first1=Éadaoin| last2=May| first2=Hila| last3=Shalem| first3=Dina| last4=Rohland| first4=Nadin| last5=Mallick| first5=Swapan| last6=Lazaridis| first6=Iosif| last7=Sarig| first7=Rachel| last8=Stewardson| first8=Kristin| last9=Nordenfelt| first9=Susanne| last10=Patterson| first10=Nick| last11=Hershkovitz| first11=Israel| last12=Reich| first12=David}}</ref>

=== Lebanon ===
 
A 2013 genetic study carried out by Haber at al found significant genetic differences between the Muslim and Christian population of Levant. According to the authors  "The population tree  splits Levantine populations in two branches: one leading to Europeans and Central Asians that includes Lebanese, Armenians, Cypriots, Druze and Jews, as well as Turks, Iranians and Caucasian populations; and a second branch composed of Palestinians, Jordanians, Syrians, as well as North Africans, Ethiopians, Saudis, and Bedouins. The tree shows a correlation between religion and the population structures in the Levant: all Jews (Sephardi and Ashkenazi) cluster in one branch; Druze from Mount Lebanon and Druze from Mount Carmel are depicted on a private branch; and Lebanese Christians form a private branch with the Christian populations of Armenia and Cyprus placing the Lebanese Muslims as an outer group. The predominantly Muslim populations of Syrians, Palestinians and Jordanians cluster on branches with other Muslim populations as distant as Morocco and Yemen." The authors explained that  "In particular, conversion of the region's populations to Islam appears to have introduced major rearrangements in populations' relations through admixture with culturally similar but geographically remote populations, leading to genetic similarities between remarkably distant populations like Jordanians, Moroccans, and Yemenis. Conversely, other populations, like Christians and Druze, became genetically isolated in the new cultural environment." In conclusions, the authors reconstructed the genetic structure of ancient Levantines and found that a pre-Islamic expansion Levant was more genetically similar to Europeans than to current Middle Easterners.<ref>{{Cite journal | doi=10.1371/journal.pgen.1003316| pmid=23468648| pmc=3585000|title = Genome-Wide Diversity in the Levant Reveals Recent Structuring by Culture| journal=PLOS Genetics| volume=9| issue=2| pages=e1003316|year = 2013|last1 = Haber|first1 = Marc| last2=Gauguier| first2=Dominique| last3=Youhanna| first3=Sonia| last4=Patterson| first4=Nick| last5=Moorjani| first5=Priya| last6=Botigué| first6=Laura R.| last7=Platt| first7=Daniel E.| last8=Matisoo-Smith| first8=Elizabeth| last9=Soria-Hernanz| first9=David F.| last10=Wells| first10=R. Spencer| last11=Bertranpetit| first11=Jaume| last12=Tyler-Smith| first12=Chris| last13=Comas| first13=David| last14=Zalloua| first14=Pierre A.}}</ref>

A 2017 genetic study carried out by Haber et al. found that Lebanon has maintained near-total genetic continuity since the Bronze Age,
which the studiers considered a surprising result considering the multiple waves of conquest. The authors of the test tested a model of the present-day Lebanese as a mixture of Sidon_BA and any other ancient Eurasian population using qpAdm, and found that the Lebanese can be best modeled as Sidon_BA 93% ± 1.6% and a Steppe Bronze Age population 7% ± 1.6%. In regards to phenotype, modern Lebanese were also found to be very close to ancient Levantine populations, with "SNPs associated with phenotypic traits showing that Sidon_BA and the Lebanese had comparable skin, hair, and eye colors (in general: light intermediate skin pigmentation, brown eyes, and dark hair) with similar frequencies of the underlying causal variants in SLC24A5 and HERC2, but with Sidon_BA probably having darker skin than Lebanese today from variants in SLC45A2 resulting in darker pigmentation."<ref>{{Cite journal | url=http://www.cell.com/ajhg/fulltext/S0002-9297(17)30276-8 | doi=10.1016/j.ajhg.2017.06.013| pmid=28757201| pmc=5544389| title=Continuity and Admixture in the Last Five Millennia of Levantine History from Ancient Canaanite and Present-Day Lebanese Genome Sequences| journal=The American Journal of Human Genetics| volume=101| issue=2| pages=274–282| year=2017| last1=Haber| first1=Marc| last2=Doumet-Serhal| first2=Claude| last3=Scheib| first3=Christiana| last4=Xue| first4=Yali| last5=Danecek| first5=Petr| last6=Mezzavilla| first6=Massimo| last7=Youhanna| first7=Sonia| last8=Martiniano| first8=Rui| last9=Prado-Martinez| first9=Javier| last10=Szpak| first10=Michał| last11=Matisoo-Smith| first11=Elizabeth| last12=Schutkowski| first12=Holger| last13=Mikulski| first13=Richard| last14=Zalloua| first14=Pierre| last15=Kivisild| first15=Toomas| last16=Tyler-Smith| first16=Chris}}</ref>

==Turkey (Anatolia)==
{{Main|Genetic history of the Turkish people}}
{{external media
| float  = right
| width  = 300px
| image1 = [https://i.redd.it/kxyqcb8fnps11.png Turic genetic impact on Asia Minor and Trace]
}}
In population genetics the question has been debated whether the modern Turkish population is significantly related to other, mainly Central Asian, Turkic peoples (from whom the modern Turkish language originates), or whether they are rather derived from the indigenous largely Indo-European and Semitic speaking ''pre-Turkic'' populations of Anatolia (Asia Minor) which were, with the exception of Greeks, Armenians, Assyrians, Jews, Georgians, Circassians and Kurds, culturally assimilated during the late Middle Ages. The contribution of the Central Asian genetics to the modern Turkish people has been debated and become the subject of several studies. As a result, several studies have concluded that the historical (pre-Islamic and pre-Turkic) and indigenous Anatolian groups are in fact the primary source of the present-day Turkish population,<ref name=antigens57>{{cite journal |last1=Arnaiz-Villena |first1=A. |last2=Karin |first2=M. |last3=Bendikuze |first3=N. |last4=Gomez-Casado |first4=E. |last5=Moscoso |first5=J. |last6=Silvera |first6=C. |last7=Oguz |first7=F.S. |last8=Sarper Diler |first8=A. |last9=De Pacho |first9=A. |last10=Allende |first10=L. |last11=Guillen |first11=J. |last12=Martinez Laso |first12=J. |date=2001 |title=HLA alleles and haplotypes in the Turkish population: Relatedness to Kurds, Armenians and other Mediterraneans |journal=Tissue Antigens |volume=57 |issue=4 |pages=308–17 |doi=10.1034/j.1399-0039.2001.057004308.x |pmid=11380939}}</ref><ref name=Yardumian_et_al>{{cite journal |doi=10.2753/AAE1061-1959500101 |last1=Schurr |first1=Theodore G. |last2=Yardumian |first2=Aram |date=2011 |title=Who Are the Anatolian Turks? |journal=Anthropology & Archeology of Eurasia |volume=50 |issue=1 |pages=6–42}}</ref><ref name="Hodoglugil_et_al2">{{cite journal |author=Hodoğlugil U |author2=Mahley RW |title=Turkish population structure and genetic ancestry reveal relatedness among Eurasian populations |journal=Annals of Human Genetics |volume=76 |issue=2 |pages=128–41 |date=March 2012  |pmid=22332727 |doi=10.1111/j.1469-1809.2011.00701.x |pmc=4904778}}</ref><ref name=eurostudy>{{cite journal |last1=Rosser |first1=Zoë H. |last2=Zerjal |first2=Tatiana |last3=Hurles |first3=Matthew E. |last4=Adojaan |first4=Maarja |last5=Alavantic |first5=Dragan |last6=Amorim |first6=António |last7=Amos |first7=William |last8=Armenteros |first8=Manuel |last9=Arroyo |first9=Eduardo |last10=Barbujani |first10=Guido |date=2000 |title=Y-Chromosomal Diversity in Europe is Clinal and Influenced Primarily by Geography, Rather than by Language |journal=The American Journal of Human Genetics |volume=67 |issue=6 |pages=1526–43 |pmid=11078479 |pmc=1287948 |doi=10.1086/316890}}</ref><ref name=humangenetics112>{{cite journal |id={{INIST|14599190}} |title=Testing hypotheses of language replacement in the neighbouring Caucasus: evidence from the Y-chromosome |journal=Human Genetics |date=2003 |volume=112 |issue=3 |pages=255–61|pmid=12596050 |author1=Nasidze |first1=I |last2=Sarkisian |first2=T |last3=Kerimov |first3=A |last4=Stoneking |first4=M |doi=10.1007/s00439-002-0874-4|url=https://link.springer.com/article/10.1007/s00439-002-0874-4 |doi-broken-date=2019-06-07 }}</ref><ref name=stanford>{{cite journal |last1=Cinnioğlu |first1=Cengiz |last2=King |first2=Roy |last3=Kivisild |first3=Toomas |last4=Kalfoğlu |first4=Ersi |last5=Atasoy |first5=Sevil |last6=Cavalleri |first6=Gianpiero L. |last7=Lillie |first7=Anita S. |last8=Roseman |first8=Charles C. |last9=Lin |first9=Alice A. |last10=Prince |first10=Kristina |last11=Oefner |first11=Peter J. |last12=Shen |first12=Peidong |last13=Semino |first13=Ornella |last14=Cavalli-Sforza |first14=L. Luca |last15=Underhill |first15=Peter A. |date=2004 |title=Excavating Y-chromosome haplotype strata in Anatolia |journal=Human Genetics |volume=114 |issue=2 |pages=127–48 |doi=10.1007/s00439-003-1031-4 |pmid=14586639}}</ref><ref name=euroasia>{{cite journal |author=Wells RS |title=The Eurasian heartland: a continental perspective on Y-chromosome diversity |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=98 |issue=18 |pages=10244–9 |date=August 2001  |pmid=11526236 |pmc=56946 |doi=10.1073/pnas.171305098 |bibcode=2001PNAS...9810244W |jstor=3056514|name-list-format=vanc |author2= Yuldasheva N |author3= Ruzibakiev R |displayauthors=3 |author4=Underhill, Peter A. |last5=Evseeva |first5=Irina |last6=Blue-Smith |first6=Jason |last7=Jin |first7=Li |last8=Su |first8=Bing |last9=Pitchappan |first9=Ramasamy |last10=Shanmugalakshmi |first10=Sadagopal |last11=Balakrishnan |first11=Karuppiah |last12=Read |first12=Mark |last13=Pearson |first13=Nathaniel M. |last14=Zerjal |first14=Tatiana |last15=Webster |first15=Matthew T. |last16=Zholoshvili |first16=Irakli |last17=Jamarjashvili |first17=Elena |last18=Gambarov |first18=Spartak |last19=Nikbin |first19=Behrouz |last20=Dostiev |first20=Ashur |last21=Aknazarov |first21=Ogonazar |last22=Zalloua |first22=Pierre |last23=Tsoy |first23=Igor |last24=Kitaev |first24=Mikhail |last25=Mirrakhimov |first25=Mirsaid |last26=Chariev |first26=Ashir |last27=Bodmer |first27=Walter F. }}</ref> in addition to neighboring peoples,<ref name="Hodoglugil_et_al2"/> such as Balkan peoples (such as Phrygians and Macedonian Greeks),<ref>{{cite journal |author=Comas D |title=Alu insertion polymorphisms in the Balkans and the origins of the Aromuns |journal=Annals of Human Genetics |volume=68 |issue=Pt 2 |pages=120–7 |date=March 2004  |pmid=15008791 |doi=10.1046/j.1529-8817.2003.00080.x|name-list-format=vanc |author2= Schmid H |author3= Braeuer S  |displayauthors=etal  }}</ref> and Central Asian Turkic people, from the Turkic homelands in modern Kazakhstan, Uzbekistan, Turkmenistan and Kirghizstan.<ref name=Hodoglugil_et_al2/>

The ''Pre-Islamic'' and ''Pre-Turkic'' speaking population of Anatolia consisted of, at different periods, a vast patchwork of Caucasoid peoples, including Language Isolate speakers such as Hurrians, Hattians and Urartians among others, who were eventually absorbed by Indo-European populations, the most prominent being Greeks, Kaskians, Persians, Luwians, Hittites, Mitanni, Phrygians, Lydians, Cimmerians, Scythians, Medes, Lycians, Cilicians, Armenians, Celts and Kurds. Semitic speaking peoples such as Assyrians, Amorites, Eblaites, Phoenicians, Arameans and Jews also maintained a longstanding presence across southern Anatolia, and to a lesser degree, Kartvelian (Georgian) and Northwest Caucasian speaking peoples in the north east.

==See also==
*Haplogroup J-P209 (Y-DNA)
*Y-DNA haplogroups in populations of the Near East
*Ancient Near East
*Genetic history of North Africa
*Genetic history of the Caucasus
*Genetic history of Europe
*Ethnic groups of the Middle East
*[[Biology:Genetic studies on Arabs|Genetic studies on Arabs]]
*[[Biology:Genetic studies on Jews|Genetic studies on Jews]]
*Genetic origins of the Turkish people
*Origin of Egyptians
*Origins of the Kurds
*Khazar theory
*Origin of the Nilotic peoples

==References==
{{Reflist|30em}}

==Bibliography==
*{{cite journal | last1 = Cruciani | first1 = F. |author2= La Fratta, R. |author3= Torroni, A. |author4= Underhill, P. A. |author5= Scozzari, R. | date = 2006 | title = Molecular Dissection of the Y Chromosome Haplogroup E-M78 (E3b1a): A Posteriori Evaluation of a Microsatellite-Network-Based Approach Through Six New Biallelic Markers | journal = Human Mutation | volume = 27  | pages = 831–2 | pmid = 16835895| doi=10.1002/humu.9445 | issue = 8 }}
*King, R. J.; Ozcan, S. S., Carter, T., Kalfoglu, E., Atasoy, S., Triantaphyllidis, C., Kouvatsi, A., Lin, A. A., Chow, C-E. T., Zhivotovsky, L. A., Michalodimitrakis, M., Underhill, P. A., (2008). "Differential Y-chromosome Anatolian Influences on the Greek and Cretan Neolithic." ''Annals of Human Genetics'' 72 Issue 2 March 2008: 205–214.
*{{cite journal | doi = 10.1002/humu.20077 | last1 = Shen | first1 = P | last2 = Lavi | first2 = T | last3 = Kivisild | first3 = T | last4 = Chou | first4 = V | last5 = Sengun | first5 = D | last6 = Gefel | first6 = D | last7 = Shpirer | first7 = I | last8 = Woolf | first8 = E | last9 = Hillel | first9 = J | date = 2004 | title = Reconstruction of Patrilineages and Matrilineages of Samaritans and Other Israeli Populations From Y-Chromosome and Mitochondrial DNA Sequence Variation | journal = Human Mutation | volume = 24 | issue = 3| pages = 248–260 | pmid = 15300852  | displayauthors = etal  }}
*Zalloua, P., Wells, S. (2004) "Who Were the Phoenicians?" ''National Geographic Magazine'', October 2004.

{{Human genetics}}

{{DEFAULTSORT:Archaeogenetics Of The Near East}}


{{Sourceattribution|Archaeogenetics of the Near East}}